Khollan Vafa : اخوان صفا: دردار، دارون، نارون، درخت پشه ( شجرالبق)، پشه غال، بوقیصا (!)، اسفیدار، بسمرقندی کل‌کرم و در صغدیان (سغدیانه*) کنجشک ( ، نام چوبی است در ساز. (غیاث ) (آنندراج ). در اندلس و در بخارا لامشکر، غرب، صنعت غرب

دردار. [ دَ ] (اِ) درختی است که پشه بار می آورد و به عربی شجرةالبق خوانند و بعضی گویند سفیددار همانست . (برهان ). آنرا دارون گویند و نام دیگرش سپیددار است و آنرا پشه دار نیز گویند. (از آنندراج ). شجرالبق خوانند و در پارسی درخت پشه خوانند و به شیرازی سفیدار. (از اختیارات بدیعی ). لغت فارسی است و او را درخت پشه و نارون نیز گویند چه ثمر او چون خشک گردد از جوف او پشه متکون می گردد، و نوعی «غرب » است . (از تحفه ٔ حکیم مؤمن ). درخت خوش سایه را گویند، و او را شجرةالبق نیز گویند. (از تذکره ٔ داود ضریر انطاکی ). قزاونه وزو خوانند، درختی بزرگ است ثمره اش مانند نار طرفی بربسته بود. (نزهة القلوب ). درختی است که پشه غال گویند و به عربی شجرالبق خوانند. (رشیدی ). شجرالبق . نارون . (بحرالجواهر). لسان العصافیر. بوقیصا*. نارون . نشم الاسود. و بیشتر پیوندی آنرا نارون و پیوندنشده ٔ آن را سیاه درخت نامند. (یادداشت مرحوم دهخدا).

* بوقیصا. (معرب ، اِ) مأخوذ از یونانی . درخت سفیدار. (ناظم الاطباء). درخت سپیدار باشد و آن نوعی از بید است و بعضی ، درخت پده و پشه غال را نیز گفته اند که بعربی شجرةالبق خوانند. (برهان ) (آنندراج ). دروار. (فهرست مخزن الادویه ) (تحفه ٔ حکیم مؤمن ). نشم الاسود . (یادداشت بخط مؤلف ). رجوع به نشم و دردار و لکلرک ج 1 ص 291 و ج 2 ص 83شود.

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نام درخت سده است که بعربی شجرةالبق خوانند. (برهان قاطع). درختی است که درون بارش پر از پشه است و بعربی شجرةالبق گویند. (فرهنگ رشیدی ). لکن این درخت را باری نیست بعضی برگهای آن ترنجیده شود و در میان آن جانورانی خرد چون پشه گرد آید. رجوع به پشه دار شود.

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دردار. شجرة البق خوانند و بپارسی درخت پشه گویند و به شیرازی اسفیدار و بسمرقندی کل‌کرم و در صغدیان کنجشک و در اندلس و در بخارا لامشکر و غرب نوعی از وی است و صنعت غرب گفته شود اما ورق دردار در وی قبضی بود و جلایی و پوست وی قابض‌تر و سردتر از ورق وی بود و طبیخ اصل وی چون بر استخوان شکسته نطول کنند سود دهد و پوست سطبر وی چون یک مثقال به آب سرد بیاشامند یا با شراب مسهل بلغم بود و ورق وی چون سحق کنند و با سرکه بر جرب ریش شده مالند نافع بود و پوست وی چون بر جراحات پیچند بصلاح آورد و اگر بکوبند و با سرکه بسرشند و بر برص طلا کنند زایل کند و بیخ وی چون بر آتش نهند و بسوزند رطوبتی که از وی بیرون آید بگیرند و در گوش چکانند کری که از رنجوری دراز حادث شده باشد زایل کند و عصاره ورق وی چون در گوش چکانند نیم‌گرم ورم گوش را نافع بود و اگر با عسل بیامیزند و در چشم کشند تاریکی چشم زایل کند و رطوبت ثمر وی چون در روی مالند جلایی تمام دهد و مسیح گوید ورق وی سرد و خشک بود در درجه اول

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صاحب مخزن الادویه می‌نویسد: دردار بفتح دال لغت فارسی است و آن را دارون و سفیدار و درخت پشه نیز نامند جهت آنکه در جوف ثمر آن پشه متکون می‌گردد و نوعی از غرب است

لاتین‌ULMUS CAMPESTRIS فرانسه‌ORME PYRAMIDL انگلیسی ‌COMMON ELM

اختیارات بدیعی، ص: 165

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نارون (نام علمی: Ulmus) نام یک سرده از تیره نارونیان است که برگریز یا نیمه‌برگریز هستند. این گونه ابتدا در دوره میوسن در حدود ۲۰ میلیون سال پیش، در جایی که امروزه آسیای مرکزی قرار دارد پدید آمد و به مرور در سراسر نیمکره شمالی گسترده شد.

گونه‌های نارون[ویرایش]

برخی از گونه‌های نارون چنین‌اند:

اوجا, Ulmus minor: این گونه در شمال ایران می‌روید.
مَلَچ, Ulmus glabra: گونه‌ای که در استان‌های گلستان، مازندران، گیلان و آذربایجان می‌روید.
نارون چتری, Ulmus Umbraculifera: گونه‌ای از نارون که در ایران و افغانستان می‌روید.
نارون برگ‌ریز یا نارون گل‌پشه‌ای, Ulmus boissieri: گونه‌ای از نارون که تنها در کشور ایران می‌روید.
نارون سفید, Ulmus laevis
نارون قرمز, Ulmus rubra

منابع[ویرایش]

مشارکت‌کنندگان ویکی‌پدیا، «Elm»، ویکی‌پدیای انگلیسی، دانشنامهٔ آزاد (بازیابی در ۲۳ فوریه ۲۰۱۴).

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به آذری و ترکی قره قاج*:

Qaraağac (lat. Ulmus) - qarağackimilər fəsiləsinə aid bitki cinsi.

Karaağaç, karaağaçgiller (Ulmaceae) familyasının Ulmus cinsinden ağaç türlerine verilen ad.(Anadolu'da karaağaç adı ile anılan ağaca,(a.Buche), kayın denilir. Bahaeddin ÖGEL, Türk Mitolojisi 2. cilt 478. sayfa, 6.Kayın Ağacı maddesi)

Dünya üzerinde ılıman iklim bölgelerinde Kuzey Amerika, Avrupa ve Asya'da yayılış gösteririler. Yapraklarınıdöken uzun ömürlü orman ağaçlarıdır. Çok sayıdaki pullu tomurcuklar sarmal dizilmiştir. Yumurtamsı konik şekilde olan yaprakların kenarları çift dişli, tabanı asimetriktir. Çiçekler erdişidir. Meyve basık endospermsizdir. Kabukları kalın ve boyuna çatlaklıdır. Kabuklar kolayca soyulur.

Diri odun açık renkli, öz odun kırmızı renklidir. odunları sert ve ağırdır. Dış darbelere dayanıklıdır. Kolay kolay kırılmaz.

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· قره قاج يا قره تاج، درختچه اي است به ارتفاع 1 تا 3 متر و بيشتر که داراي برگ هايي مرکب از 3 برگچه با دمبرگ دراز است.
گياه سرسبز «قره قاج» که در شهرستان زيباي گيلا نغرب، به وفور يافت مي شود، از خانواده «بقولا ت» يا «minoseae Leg» داراي 3 زيرخانواده است که «Anagyrigsfoetiba» به زير خانواده «Papiliononoi» يا پروانه واران (پروانه آسا) متعلق هست. (البته بعضي از گياه شناسان، اين سه زير خانواده را، هر کدام يک خانواده مستقل مي دانند، که در اين صورت، درختچه «Anagarisfoetibe» جزو خانواده «Plionaceae P» محسوب مي شود).
خانواده بقولا ت، بعد از خانواده کاسني بزرگترين خانواده گياهي هستند که شامل 55 جنس و حدود 13000 گونه هستند و انتشار جهاني دارند و در حدود 140 جنس از اين تيره به عنوان درختچه هاي زينتي کاشته مي شوند.
نام علمي درختچه براي اولين بار توسط «لينه» (1778-1707) م- گياه شناس معروف قرن 18 ميلا دي و پدر علم رده بندي گياهي، به کار برده شد.
«Anagyrigsfoetiba» حدود 375 جنس را در بر مي گيرد و به 10 طايفه تقسيم مي شوند که بيشتر در مناطق معتدله جنوبي و شمالي انتشار دارند. از نظر رده بندي گياهي ، زير خانواده پروانه واران، از دو زير خانواده ديگر، تکامل يافته ترند.
ويژگي هاي مرفولوژيکي گياه: علا وه بر اينکه اين گياهان معمولا از ارتفاعي بين 1 تا 3 متر بهره مندند، داراي تخمدان فوقاني يک حجره اي نيز هستند. از مشخصات ديگر گياه قره قاج، داشتن کاسبرگ به هم چسبيده که در انتها آزاد هستند، مي باشد.
گلبرگ که خارجي ترين عضو گياه به شمار مي رود، در پشت قرار گرفته و دو تار داخلي دارد که به آن بال مي گويند و دو تار ديگر داخلي ترين هستند که به آن ناو مي گويند که غالبا بهم متصل و پرچم و مادگي را در بر مي گيرد. گل هاي قره قاج، که به نوعي در شهرستان معتدل گيلا نغرب، زبانزد خاص و عام است به رنگ زرد و ميوه اش به صورت نيام (شمشير)، قوسي شکل، به درازاي10 تا 15 سانتيمتر و منتهي به نوک باريک و دراز است. در داخل ميوه آن 3 تا 8 دانه به رنگ بنفش و شبيه دانه لوبيا جاي دارد که اهالي شهرستان معتدل گيلا نغرب به همين دليل به آن شبدر لوبيا مي گويند. دانه اين گياه خوش رنگ، آنقدر سخت است که ممکن است مدت ها در خاک بماند ولي سبز نشود. معمولا براي سبز شدن روي آن را خراش داده يا اسيد کربنيک مي ريزند. از تمام قسمت هاي گياه با طراوت قره قاج، بوي نامطبوع استشمام مي شود. دانه آن داراي اثر سمي است و خطر آن نيز در اين است که اطفال آن را اشتباها با دانه لوبيا، مخلوط کنند. با مصرف دانه آن استفراغ هاي منجر به حالت خوني پيش آمده است و اين مورد يکي از خواص بي شمار دارويي قره قاج است که تنها، دانه آن، قي آور است.
شير حيواناتي که گياه زينتي قره قاج، مصرف کنند نيز بوي ناپسند دارد.
ترکيبات شيميايي گياه قره قاج: در اعضاي مختلف اين گياه 2 الکالوئيد سمي، يکي به نام «سيتيزين Cytisine» و ديگري «آناژيرين Anagyrine» وجود دارد. الکالوئيد اخير که توسط «هاردي و allois در سال 1885 م از گياه مذکور به دست آمد، در اندام هاي مختلف گياه، مخصوصا در برگ و پارانشيم اعضاي مسن، اشعه مغزي در بين آوندهاي آبکش و بيشتر از همه در بين لپه هاي دانه يافت مي شود. در دانه اين گياه علا وه بر آناژيرين، ماده روغني و يک ماده ملون به رنگ زرد نيز وجود دارد.

اکوسيستم گياه وحشي قره قاج

شرايط آب و هوايي: اين درختچه زيبا، از نظر اقليمي در مناطق آب و هواي معتدل مديترانه اي و معمولا بين مرز دو اکوسيستم گرمسيري و سردسيري، موسوم به اکوتون ديده مي شود. شهرستان مديترانه اي «گيلا نغرب» نمود بارز اين شرايط آب و هوايي است که ميزبان جنگل سبز «قره قاج» بوده و هست. محدوده مورد نظر، به دليل واقع شدن در کوهستان سترگ زاگرس و محل عبور سيکلون هاي غربي، داراي بارش بين 600-500 ميليمتر است. درختچه زيباي قره قاج، در منطقه زمين لغزه «کاسه کران» روستايي ديدني، از توابع دهستان سردسير «چله» گيلا نغرب که مرز بين دو اکوسيستم گرم گيلا نغرب و اکوسيستم محسوب مي شود ديده شده است.
زمين شناسي منطقه: ناحيه مورد نظر از بعد ژئومورفولوژي، يک زمين لغزه معروف مي باشد که لا يه آهک آسماري با ضخامت حدود 100 متر، بر روي لا يه «مارني آهکي سازند پابده» لغزيده و سطح گسيختگي به مساحت 9/5 کيلومتر مربع و نيز توده لغزيده از سنگ و خاک به وسعت 13/5 کيلومتر مربع به وجود آمده است. اين بهم ريختگي باعث نفوذپذيري نسبتا خوب خاک شده و پوشش جنگلي بسيار متراکم، خصوصا جنگل «قره قاج» در دامنه کوهستان را موجب شده است. در قسمت هايي از اين منطقه که سطح بالا ي سازند مارني فرسايش يافته، به علت نرمي و فرسايش پذيري شديد مارن دره هاي عميق ناشي از فرسايش آبي به وجود آمده که متاسفانه به شکل لکه هاي سفيد بدون درخت، در ميان انبوهي از درختان کمياب «بلوط» و «قره قاج» ديده مي شود که تهديدي فوق العاده جدي براي اکوسيستم منطقه است.
خاک هاي رسوبي و آبرفتي، با آهک و مارن در اثر ناپايداري و لغزش با هم مخلوط شده و در بعضي جاها نيز حالت برش پي درپي به وجود آمده است.
اين تنوع خاک توانسته تنوع زيستي خاصي را به وجود آورد که وجود درختان متراکم بلوط نادر و قره قاج کمياب و بوته هاي خاردار، حکايت از اين پديده دارد.

اثرات اکوسيستم «قره قاج»

اين درختچه نادر به دليل ويژگي هاي متفاوتي که با ساير پوشش گياهي دارد، تنوع زيستي خاصي را در منطقه موجب شده است که از اين قرارند:
1- اعتدال آب و هوا: اين گونه کمياب به دليل تنوع در دوره رشد آن، خصوصيات ويژه اي را از نظر اقليمي به وجود آورده است. چنانکه اواخر پائيز در نهايت شگفتي و تفکر در اسرار بقا، «قره قاج» جوانه مي زند و تقريبا تا اواسط تيرماه، سبز است. اين در حالي است که اکثر نباتات با شروع فصل خزان علا وه بر گرايش برگ ها به رنگ هاي زرد، ارغواني و قهوه اي و سرانجام ريزش آنها و ورود به زمان خواب زمستاني، در حقيقت چهره سبز طبيعت را بي رنگ مي کنند.
2- حالت تزئيني بودن «قره قاج»: حالتي است که مقارن تجربه کردن درختان در خواب زمستاني، با شکوفا شدن گل هاي زرد و سفيد، چنان زيبايي خاصي به منطقه مي بخشد که گويي بهار، در زمستان خلق شده است و اين مورد به خصوص، يکي از شاهکارهاي طبيعت شهرستان زيباي «گيلا نغرب» است که در طول سال 2 بار بهار را تجربه مي کند.
3- حفاظت خاک: به دليل تراکم زياد گياه کمياب «قره قاج» مخصوصا در شاخ و برگ، از شدت بارش به سطح زمين، به نحو چشمگيري جلوگيري کرده و آب به آرامي در پاي درختچه به زمين فرو مي رود که هم موجب نفوذپذيري بيشتر مي شود و هم اينکه خاک فرسايش نمي يابد.
بنابراين وجود رطوبت در پاي «قره قاج» باعث رشد گياهان فراوان در همسايگي درختچه مي شود.
4- حفاظت از جانوران در مقابل سرماي زمستان و گرماي اواخر بهار: درختچه هاي زيبا و بسيار «پر اهميت» «قره قاج» در اکثر مواقع آنچنان متراکم اند، که عبور و مرور دام ها و نيز افراد، در داخل آنها، به راحتي مقدور نمي باشد. به همين دليل پناهگاه خوبي براي وحوش نادر منطقه که جزو ذخاير طبيعي کشور به شمار مي روند، محسوب مي شوند که اين مهم در اکوسيستم کم نظير «شهرستان زيبا و مديترانه اي گيلا نغرب» بايد جدي تلقي شود.
خصوصا آنکه تعدادي از جانداران، مي توانند در تمام طول سال، حتي در تابستان که «قره قاج» لباس سبز خود را از تن بدر مي کند، در لا به لا ي اين گونه هاي ارزشمند گياهي، مخصوصا براي جانوران، علا وه بر تداوم حيات، از خطرات در امان بمانند.
5- خواص دارويي «قره قاج»: برگ، ساقه و ريشه اين گياه مقدس، در درمان بعضي از امراض، حکم کيميا را دارد. چنانکه اثر مسهل بودن و ضد انگل آن و مصرف مقادير زياد آن، قي آور و قاعده آور است. اثر دانه گياه، قوي تر از اعضاي ديگر آن است. له شده برگ گياه تازه، را به صورت «ضماد» بر روي «زخم ها» و «تومورها»، جهت بهبود آنها قرار مي دهند و نيز دم کرده (8 تا 16 گرم) ساقه، برگ و ريشه، مسهل خوبي مي تواند باشد.

موقعيت

اين درختچه به صورت جنگلي، در حوزه آبخيز «دهستان سردسير چله»، از توابع «گيلا نغرب»، از روستاي «کاسه کران» تا منطقه «تق توق»، در محل زمين لغزه بزرگ «کاسه کران» رويش داشته است. زمين لغزه مزبور، که اينک اکوسيستمي از گونه نادر، دارويي، زيبا، با طبيعت عجيب را در خود جا داده در زمان هاي بسيار دور، رخ داده است، به طوري که امروزه اثرات اين لغزش بزرگ به خوبي معلوم است. عرض جغرافيايي آن (34/7-34/4) و طول (46/17 -6/14) شرقي، در 5 کيلومتري شهرستان گيلا نغرب، واقع شده است.
اين محدوده در دامنه شمال تا قديس «سريوان» به مساحت تقريبي 900 هکتار که منطبق بر زمين لغزش کاسه کران است رويش دارد.
6- جهت تغذيه دام: قره قاج، خوراک مناسبي براي چراي دام ها مي باشد (و اين نکته حائز اهميت است که شهرستان گيلا نغرب هرساله ميزبان صدها هزار راس گوسفند جهت تعليف مي باشد) و اين مورد ويژه، اين را مي رساند که گياه مزبور مي تواند در مناطق ديگر شهرستان و جاهايي که آب و هوايي شبيه آب و هواي گيلا نغرب دارد، کاشته شود. حتي در گلخانه ها.
7- برگزاري اردوهاي علمي و توريستي: «قره قاج» و اصولا جنگل سرسبز، با طراوت، کمياب و بسيار سودمند آن، اين استعداد و پتانسيل را دارا است که هر ساله، ميزبان تورهاي علمي و نيز گردشگري از داخل و خارج کشور باشد. مغناطيس گياهي که مخصوصا طرفداران محيط زيست و گونه هاي مهم نباتي جهان را به سوي خود جذب مي کند و چشم هر رهگذري را نوازش مي دهد و اين همت مسوولين را مي طلبد.

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به عربی دردار:

الدردار أو البوقيصا^[1] (بالإنجليزية: Elm) (الاسم العلمي:Ulmus) هو جنس نباتي يتبع فصيلة الدردارية من رتبة الورديات.^[2][3][4][5] وهو جنس من الأشجار النفضية ظهرت في الميوسين منذ نحو 40 مليون سنة في ما يقابل آسيا الوسطى المعاصرة.

محتويات

[أظهر]

من أنواعه في الوطن العربي[عدل]

الدردار الأجرد (باللاتينية: Ulmus glabra) في بلاد الشام وأوروبا
الدردار الأصغر (باللاتينية: Ulmus minor) ويتبعه الدردار الأصغر نويع الرمادي (باللاتينية: Ulmus minor subsp. canescens) في بلاد الشام والمغرب العربي وبعض مناطق أوروبا
الدردار الطويل (باللاتينية: Ulmus procera) في المغرب العربي والبلقان وبعض مناطق أوروبا

من أنواعه الأخرى[عدل]

الدردار الأحمر (باللاتينية: Ulmus rubra)
الدردار الإسماعيلي (باللاتينية: Ulmus ismaelis)
الدردار التوماسي (باللاتينية: Ulmus thomasii)
الدردار الخفيف (باللاتينية: Ulmus laevis)
الدردار الصغير (باللاتينية: Ulmus nana)
الدردار صغير الثمار (باللاتينية: Ulmus microcarpa)
الدردار العالي (باللاتينية: Ulmus alata)
الدردار كبير الثمار (باللاتينية: Ulmus macrocarpa)
الدردار المتطاول (باللاتينية: Ulmus elongata)

أنظر أيضاً[عدل]

قائمة أنواع الدردار

مراجع[عدل]

^ موسوعة شبكة المعرفة الريفية. الدردار. تاريخ الولوج 30 أيلول 2011.
^ موقع لائحة النباتات أنواع الدردار تاريخ الولوج 16 ديسمبر 2012
^ موقع زيبكودزو جنس الدردار تاريخ الولوج 16 ديسمبر 2012
^ المركز الوطني للمعلومات التقنية الحيوية جنس الدردار تاريخ الولوج 16 ديسمبر 2012
^ موقع تاكسونوميكون جنس الدردار تاريخ الولوج 16 ديسمبر 2012

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به عبری اُلموس:

אולמוס או בוקיצה (שם מדעי: Ulmus) הוא עץ נשיר ממשפחת המישיים. ישנם בין 30 ל-40 מינים של העץ, תשעה מהם ניתן למצוא בצפון אמריקה וכן מספר קטן יותר באירופה. את המגוון הגדול ביותר ניתן למצוא בסין. בישראל צומח עץ ממשפחת האולמוס בשם בוקיצה שעירה(לשעבר: אולמוס שעיר).

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Elms are deciduous and semi-deciduous trees comprising the flowering plant genus Ulmus in the plant familyUlmaceae. The genus first appeared in the Miocene geological period about 20 million years ago, originating in what is now central Asia.^[1] These trees flourished and spread over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North America and Eurasia, presently ranging southward across the Equator into Indonesia.

Elms are components of many kinds of natural forests. Moreover, during the 19th and early 20th centuries many species and cultivars were also planted as ornamental street, garden, and park trees in Europe, North America, and parts of the Southern Hemisphere, notably Australasia. Some individual elms reached great size and age. However, in recent decades, most mature elms of European or North American origin have died from Dutch elm disease, caused by a microfungus dispersed by bark beetles. In response, disease-resistant cultivars have been developed, capable of restoring the elm to forestry and landscaping.

Contents

[show]

Taxonomy[edit]

Main articles: List of Elm species and varieties by common name, Alphabetical list of extant Elm species by scientific name, List of Elm species and varieties by scientific name, List of Elm cultivars, hybrids and hybrid cultivars and List of Elm synonyms and accepted names

There are about 30 to 40 species of Ulmus (elm); the ambiguity in number results from difficulty in delineating species, owing to the ease of hybridization between them and the development of local seed-sterile vegetatively propagated microspecies in some areas, mainly in the field elm (Ulmus minor) group. Rackham^[2] describes Ulmusas the most difficult critical genus in the entire British flora, adding that 'species and varieties are a distinction in the human mind rather than a measured degree of genetic variation'. Eight species are endemic to North America, and a smaller number to Europe;^[3] the greatest diversity is found in Asia.^[4]

The classification adopted in the List of elm species, varieties, cultivars and hybrids is largely based on that established by Brummitt.^[5] A large number of synonyms have accumulated over the last three centuries; their currently accepted names can be found in the list List of elm Synonyms and Accepted Names.

Botanists who study elms and argue over elm identification and classification are called pteleologists, from the Greek πτελέα (:elm).^[6]

As part of the sub-order urticalean rosids they are distant cousins of cannabis, hops, and nettles.

Etymology[edit]

The name Ulmus is the classical name for these trees, with the English name "elm" and many other European names derived from it.^[7]

Description[edit]

The genus is hermaphroditic, having apetalous perfect flowers which are wind-pollinated. Elm leaves are alternate, with simple, single- or, most commonly, doubly serrate margins, usually asymmetric at the base and acuminate at the apex. The fruit is a round wind-dispersed samara flushed with chlorophyll, facilitating photosynthesis before the leaves emerge.^[8] All species are tolerant of a wide range of soils and pH levels but, with few exceptions, demand good drainage.

'Sapporo Autumn Gold', Antella, Florence

Wych elm (Ulmus glabra) leaves and seeds

Asymmetry of leaf, Slippery Elm U. rubra

Mature bark, Slippery Elm U. rubra

Flowers of the hybrid elm cultivar 'Columella'

Corky wings, Winged Elm U. alata

U. laciniata samara

U. americana, Dufferin St., Toronto, c. 1914

Pests and diseases[edit]

Main article: List of elm diseases

Dutch elm disease[edit]

Main article: Dutch elm disease

Dutch elm disease (DED) devastated elms throughout Europe and much of North America in the second half of the 20th century. It derives its name 'Dutch' from the first description of the disease and its cause in the 1920s by the Dutch botanists Bea Schwarz and Christina Johanna Buisman. Owing to its geographical isolation and effective quarantine enforcement, Australia has so far remained unaffected by Dutch Elm Disease, as have the provinces ofAlberta and British Columbia in western Canada.

DED is caused by a micro-fungus transmitted by two species of Scolytus elm-bark beetle which act as vectors. The disease affects all species of elm native to North America and Europe, but many Asiatic species have evolved anti-fungal genes and are resistant. Fungal spores, introduced into wounds in the tree caused by the beetles, invade the xylem or vascular system. The tree responds by producing tyloses, effectively blocking the flow from roots to leaves. Woodland trees in North America are not quite as susceptible to the disease because they usually lack the root-grafting of the urban elms and are somewhat more isolated from each other. In France, inoculation with the fungus of over three hundred clones of the European species failed to find a single variety possessed of any significant resistance.

The first, less aggressive strain of the disease fungus, Ophiostoma ulmi, arrived in Europe from the Far East in 1910, and was accidentally introduced to North America in 1928, but was steadily weakened by viruses and had all but disappeared in Europe by the 1940s. The second, far more virulent strain of the disease Ophiostoma novo-ulmi was identified in Europe in the late 1960s, and within a decade had killed over 20 million trees (approximately 75%) in the UK alone. Approximately three times more deadly, the new strain arrived in Europe from the USA on a cargo of Rock Elm; the hypothesis that it arose from a hybrid between the original O. ulmi and another strain endemic to the Himalaya, Ophiostoma himal-ulmi is now discredited.^[9]

There is no sign of the current pandemic waning, and no evidence of a susceptibility of the fungus to a disease of its own caused by d-factors: naturally occurring virus-like agents that severely debilitated the original O. ulmi and reduced its sporulation.^[10]

Elm phloem necrosis[edit]

Elm phloem necrosis (elm yellows) is a disease of elm trees that is spread by leafhoppers or by root grafts.^[11] This very aggressive disease, with no known cure, occurs in the Eastern United States, southern Ontario in Canada, and Europe. It is caused by phytoplasmas which infect the phloem (inner bark) of the tree.^[12] Infection and death of the phloem effectively girdles the tree and stops the flow of water and nutrients. The disease affects both wild-growing and cultivated trees. Occasionally, cutting the infected tree before the disease completely establishes itself and cleanup and prompt disposal of infected matter has resulted in the plant's survival via stump-sprouts.

Insects[edit]

Most serious of the elm pests is the elm leaf beetle Xanthogaleruca luteola, which can decimate foliage, although rarely with fatal results. The beetle was accidentally introduced to North America from Europe. Another unwelcome immigrant to North America is the Japanese beetle Popillia japonica. In both instances the beetles cause far more damage in North America owing to the absence of the predators present in their native lands. In Australia, introduced elm trees are sometimes used as foodplants by the larvae of hepialid moths of the genus Aenetus. These burrow horizontally into the trunk then vertically down.^[13][14]

Birds[edit]

Sapsucker woodpeckers have a great love of young elm trees.

Development of trees resistant to Dutch elm disease[edit]

See also: List of Elm cultivars, hybrids and hybrid cultivars

Efforts to develop resistant cultivars began in the Netherlands in 1928 and continued, uninterrupted by World War II, until 1992.^[15] Similar programmes were initiated in North America (1937), Italy (1978), and Spain (1990s). Research has followed two paths:

Species and species cultivars[edit]

In North America, careful selection has produced a number of trees resistant not only to disease, but also to the droughts and extremely cold winters afflicting the continent. Research in the USA has concentrated on the American Elm U. americana, resulting in the release of highly resistant clones, notably the cultivars 'Valley Forge' and 'Jefferson'. Much work has also been done into the selection of disease-resistant Asiatic species and cultivars.^[16][17]

In Europe, the European White Elm Ulmus laevis has received much attention. Whilst this elm has little innate resistance to Dutch elm disease, it is not favoured by the vector bark beetles and thus only becomes colonized and infected when there are no other choices, a rare situation in western Europe. Research in Spain has suggested that it may be the presence of a triterpene, alnulin, which makes the tree bark unattractive to the beetle species that spread the disease.^[18] However this possibility has not been conclusively proven.^[19] More recently, Field Elms Ulmus minor highly resistant to DED have been discovered in Spain, and form the basis of a major breeding programme.^[20]

Hybrid cultivars[edit]

Owing to their innate resistance to Dutch elm disease, Asiatic species have been crossed with European species, or with other Asiatic elms, to produce trees which are both highly resistant to disease and tolerant of native climates. After a number of false dawns in the 1970s, this approach has produced a range of reliable hybrid cultivars now commercially available in North America and Europe.^{[21][22][23][24][25][26][27]}

However, some of these cultivars, notably those with the Siberian Elm U. pumila in their ancestry, lack the forms for which the iconic American Elm and English Elm were prized. Moreover, several exported to northwestern Europe have proven unsuited to the maritime climate conditions there, notably because of their intolerance of anoxic conditions resulting from ponding on poorly drained soils in winter. Dutch hybridizations invariably included theHimalayan Elm U. wallichiana as a source of anti-fungal genes and have proven more tolerant of wet ground; they should also ultimately reach a greater size. However, the susceptibility of the cultivar 'Lobel', used as a control in Italian trials, to elm yellows has now (2014) raised a question mark over all the Dutch clones.^[28]

A number of highly resistant Ulmus cultivars has been released since 2000 by the Institute of Plant Protection in Florence, most commonly featuring crosses of the Dutch cultivar 'Plantijn' with the Siberian Elm to produce resistant trees better adapted to the Mediterranean climate.^[22]

Cautions regarding novel cultivars[edit]

Elms take many decades to grow to maturity, and as the introduction of these disease-resistant cultivars is relatively recent, their long-term performance and ultimate size and form cannot be predicted with certainty. The National Elm Trial in North America, begun in 2005, is a nationwide trial to assess strengths and weaknesses of the 19 leading cultivars raised in the US over a ten-year period; European cultivars have been excluded. Meanwhile, in Europe, American and European cultivars are being assessed in field trials started in 2000 by the UK charity Butterfly Conservation.^[29]

The ball-headed graft narvan elm, Ulmus minor 'Umbraculifera', cultivated in Persia and widely planted in central Asia.

Lafayette Street, Salem, Massachusetts: an example of the 'high-tunnelled effects' of Ulmus americana avenues once common in New England

Uses of elms in landscaping[edit]

Camperdown elm (Ulmus'Camperdown'), cultivated in Prospect Park, Brooklyn, New York

An avenue of elm trees in Fitzroy Gardens, Melbourne

One of the earliest of ornamental elms was the ball-headed graft narvan elm, Ulmus minor 'Umbraculifera', cultivated from time immemorial in Persia as a shade tree and widely planted in cities through much of south-west and central Asia. From the 18th century to the early 20th century, elms, whether species, hybrids or cultivars, were among the most widely planted ornamental trees in both Europe and North America. They were particularly popular as a street tree in avenue plantings in towns and cities, creating high-tunnelled effects. Their quick growth and variety of foliage and forms,^[30] their tolerance of air-pollution and the comparatively rapid decomposition of their leaf-litter in the fall were further advantages. In North America, the species most commonly planted was the American elm (Ulmus americana), which had unique properties that made it ideal for such use: rapid growth, adaptation to a broad range ofclimates and soils, strong wood, resistance to wind damage, and vase-like growth habit requiring minimal pruning. In Europe, the wych elm (Ulmus glabra) and the Field Elm (Ulmus minor) were the most widely planted in the countryside, the former in northern areas including Scandinavia and northern Britain, the latter further south. The hybrid between these two, Dutch elm (U. × hollandica), occurs naturally and was also commonly planted. In much of England, it was the English Elm which later came to dominate the horticultural landscape. Most commonly planted in hedgerows, it sometimes occurred in densities of over 1000 per square kilometre. In south-eastern Australia and New Zealand, large numbers of English and Dutch elms, as well as other species and cultivars, were planted as ornamentals following their introduction in the 19th century, while in northern Japan Japanese Elm (Ulmus davidiana var. japonica) was widely planted as a street tree. From about 1850 to 1920, the most prized small ornamental elm in parks and gardens was the Camperdown elm (Ulmus glabra 'Camperdownii'), a contorted weeping cultivar of the Wych Elm grafted on to a non-weeping elm trunk to give a wide, spreading and weeping fountain shape in large garden spaces.

In northern Europe elms were, moreover, among the few trees tolerant of saline deposits from sea spray, which can cause "salt-burning" and die-back. This tolerance made elms reliable both as shelterbelt trees exposed to sea wind, in particular along the coastlines of southern and western Britain^[31][32] and in the Low Countries, and as trees for coastal towns and cities.^[33]

This belle époque lasted until the First World War, when as a consequence of hostilities, notably in Germanywhence at least 40 cultivars originated, and of the outbreak at about the same time of the early strain of Dutch elm disease, Ophiostoma ulmi, the elm began its slide into horticultural decline. The devastation caused by the Second World War, and the demise in 1944 of the huge Späth nursery in Berlin, only accelerated the process. The outbreak of the new, three times more virulent, strain of Dutch elm disease Ophiostoma novo-ulmi in the late 1960s brought the tree to its nadir.

Since circa 1990 the elm has enjoyed a renaissance through the successful development in North America and Europe of cultivars highly resistant to the new disease.^[8] Consequently, the total number of named cultivars, ancient and modern, now exceeds 300, although many of the older clones, possibly over 120, have been lost to cultivation. Some of the latter, however, were by today's standards inadequately described or illustrated before the pandemic, and it is possible that a number survive, or have regenerated, unrecognised. Enthusiasm for the newer clones often remains low owing to the poor performance of earlier, supposedly disease-resistant Dutch trees released in the 1960s and 1970s. In the Netherlands, sales of elm cultivars slumped from over 56,000 in 1989 to just 6,800 in 2004,^[34] whilst in the UK, only four of the new American and European releases were commercially available in 2008.

Other uses of elms[edit]

Wood[edit]

Elm wood

Elm in boat-building: John Constable, Boat-building near Flatford Mill, 1815 (landscape with hybrid elmsUlmus × hollandica^[1])

English longbow of elm

Elm wood is valued for its interlocking grain, and consequent resistance to splitting, with significant uses in wagonwheel hubs, chair seats and coffins. The bodies of Japanese Taiko drums are often cut from the wood of old elm trees, as the wood's resistance to splitting is highly desired for nailing the skins to them, and a set of three or more is often cut from the same tree. The elm's wood bends well and distorts easily making it quite pliant. The often long, straight, trunks were favoured as a source of timber for keels in ship construction. Elm is also prized by bowyers; of the ancient bows found in Europe, a large portion of them are elm. During the Middle Ages elm was also used to make longbows if yew was unavailable.

The first written references to elm occur in the Linear B lists of military equipment at Knossos in the Mycenaean Period. Several of the chariots are of elm (« πτε-ρε-ϝα », pte-re-wa), and the lists twice mention wheels of elmwood.^[35] Hesiod says that ploughs in Ancient Greece were also made partly of elm.^[36]

The density of elm wood varies between species, but averages around 560 kg per cubic metre.^[37]

Elm wood is also resistant to decay when permanently wet, and hollowed trunks were widely used as water pipes during the medieval period in Europe. Elm was also used as piers in the construction of the original London Bridge. However this resistance to decay in water does not extend to ground contact.^[37]

Viticulture[edit]

The Romans, and more recently the Italians, used to plant elms in vineyards as supports for vines. Lopped at three metres, the elms' quick growth, twiggy lateral branches, light shade and root-suckering made them ideal trees for this purpose. The lopped branches were used for fodder and firewood.^[38] Ovid in his Amores characterizes the elm as "loving the vine": ulmus amat vitem, vitis non deserit ulmum (:the elm loves the vine, the vine does not desert the elm),^[39] and the ancients spoke of the "marriage" between elm and vine.^[40]

Medicinal products[edit]

The mucilaginous inner bark of the Slippery Elm Ulmus rubra has long been used as a demulcent, and is still produced commercially for this purpose in the United States with approval for sale as a nutritional supplement by the U.S. Food and Drug Administration.^[41]

Fodder[edit]

Elms also have a long history of cultivation for fodder, with the leafy branches cut to feed livestock. The practice continues today in the Himalaya, where it contributes to serious deforestation.^[42]

Biomass[edit]

As fossil fuel resources diminish, increasing attention is being paid to trees as sources of energy. In Italy, the Istituto per la Protezione delle Piante is (2012) in the process of releasing to commerce very fast-growing elm cultivars, able to increase in height by more than 2 m (6 ft) per annum.^[43]

Food[edit]

Elm bark, cut into strips and boiled, sustained much of the rural population of Norway during the great famine of 1812. The seeds are particularly nutritious, containing 45% crude protein, and less than 7% fibre by dry mass.^[44]

Internal mill-wheel of elm, De Hoop mill, Oldebroek, Netherlands

Alternative medicine[edit]

Elm has been listed as one of the 38 substances that are used to prepare Bach flower remedies,^[45] a kind ofalternative medicine promoted for its effect on health. However, according to Cancer Research UK, "there is no scientific evidence to prove that flower remedies can control, cure or prevent any type of disease, including cancer".^[46]

Bonsai[edit]

Chinese Elm Ulmus parvifoliabonsai

Chinese elm Ulmus parvifolia is a popular choice for bonsai owing to its tolerance of severe pruning.

Genetic resource conservation[edit]

In 1997, a European Union elm project was initiated, its aim to coordinate the conservation of all the elm genetic resources of the member states and, among other things, to assess their resistance to Dutch elm disease. Accordingly, over 300 clones were selected and propagated for testing.^[47][48][49]

Notable elm trees[edit]

Main article: List of notable elm trees

Many elm (Ulmus) trees of various kinds have attained great size or otherwise become particularly noteworthy.

The elm in art[edit]

Many artists have admired elms for the ease and grace of their branching and foliage, and have painted them with sensitivity. Elms are a recurring element in the landscapes and studies of, for example, John Constable, Ferdinand Georg Waldmüller, Frederick Childe Hassam, Karel Klinkenberg,^[50] and George Inness.

John Constable, 'Study of an Elm Tree' [1821]

John Constable, 'The Cornfield' [1826] (Ulmus × hollandica^[1])

Constable, 'Salisbury Cathedral from the Bishop's Garden' [1823 version] (Ulmus × hollandica^[1])

Jacob George Strutt,Elms at Mongewell, Oxfordshire [1830] (Ulmus minor 'Atinia')

Ferdinand Georg Waldmüller, 'Alte Ulmen im Prater' (:Old Elms inPrater) [1831]

James Duffield Harding, 'The Great Exhibition of 1851' (Ulmus minor'Atinia', centre)

Arthur Hughes, 'Home from Sea' [1862] (Ulmus minor 'Atinia' ^[1])

Ford Madox Brown, 'Work' [1863] (Ulmus minor 'Atinia' ^[1])

[unknown artist] The American Elm [1879] (Ulmus americana)

Johannes Karel Christiaan Klinkenberg, 'Amsterdam' [1890] (Ulmus x hollandica‘Belgica’ )

Frederick Childe Hassam, 'Champs Elysées, Paris' [1889] (Ulmus × hollandica, 'orme femelle'^[1])

Frederick Childe Hassam, 'Washington Arch, Spring' [1893] (Ulmus americana)

Frederick Childe Hassam, 'Church at Old Lyme' [1905] (Ulmus americana)

Frederick Childe Hassam, 'The East Hampton Elms in May' [1920] (Ulmus americana)

George Inness, 'Old Elm at Medfield' (Ulmus americana)

Unknown artist, 'The Cam near Trinity College, Cambridge', England (Ulmus atinia)

The elm in mythology and literature[edit]

Achilles and Scamander

Dryad

In Greek mythology the nymph Ptelea (Πτελέα, Elm) was one of the eight Hamadryads, nymphs of the forest and daughters of Oxylos and Hamadryas.^[51]

The first reference in literature to elms occurs in the Iliad. When Eetion, father ofAndromache, is killed by Achilles during the Trojan War, the Mountain Nymphs plant elms on his tomb («περὶ δὲ πτελέoι εφύτεψαν νύμφαι ὀρεστιάδες, κoῦραι Διὸς αἰγιόχoιo»).^[52]Also in the Iliad, when the River Scamander, indignant at the sight of so many corpses in his water, overflows and threatens to drown Achilles, the latter grasps a branch of a great elm in an attempt to save himself («ὁ δὲ πτελέην ἕλε χερσὶν εὐφυέα μεγάλην».^[53]

The Nymphs also planted elms on the tomb in the Thracian Chersonese of "great-heartedProtesilaus’’ («μεγάθυμου Πρωτεσιλάου»), the first Greek to fall in the Trojan War. These elms grew to be the tallest in the known world; but when their topmost branches saw far off the ruins of Troy, they immediately withered, so great still was the bitterness of the hero buried below, who had been loved byLaodamia and slain by Hector.^[54][55][56] The story is the subject of a poem by Antiphilus of Byzantium (1st century AD) in the Palatine Anthology:

Θεσσαλὲ Πρωτεσίλαε, σὲ μὲν πολὺς ᾄσεται αἰών,

Tρoίᾳ ὀφειλoμένoυ πτώματος ἀρξάμενoν•

σᾶμα δὲ τοι πτελέῃσι συνηρεφὲς ἀμφικoμεῦση

Nύμφαι, ἀπεχθoμένης Ἰλίoυ ἀντιπέρας.

Δένδρα δὲ δυσμήνιτα, καὶ ἤν ποτε τεῖχoς ἴδωσι

Tρώϊον, αὐαλέην φυλλοχoεῦντι κόμην.

ὅσσoς ἐν ἡρώεσσι τότ᾽ ἦν χόλoς, oὗ μέρoς ἀκμὴν

ἐχθρὸν ἐν ἀψύχoις σώζεται ἀκρέμoσιν.^[57]

[:Thessalian Protesilaos, a long age shall sing your praises,

Of the destined dead at Troy the first;

Your tomb with thick-foliaged elms they covered,

The nymphs, across the water from hated Ilion.

Trees full of anger; and whenever that wall they see,

Of Troy, the leaves in their upper crown wither and fall.

So great in the heroes was the bitterness then, some of which still

Remembers, hostile, in the soulless upper branches.]

Protesilaus had been king of Pteleos (Πτελεός) in Thessaly, which took its name from the abundant elms (πτελέoι) in the region.^[58]

Elms occur often in Pastoral Poetry, where they symbolise the idyllic life, their shade being mentioned as a place of special coolness and peace. In the first Idyll of Theocritus, for example, the goat-herd invites the shepherd to sit "here beneath the elm" («δεῦρ’ ὑπὸ τὰν πτελέαν») and sing. Beside elms Theocritus places "the sacred water" («το ἱερὸν ὕδωρ») of the Springs of the Nymphs and the shrines to the nymphs.^[59]

Aside from references literal and metaphorical to the elm and vine theme, the tree occurs in Latin literature in the Elm of Dreams in the Aeneid.^[60] When the Sibyl of Cumae leads Aeneas down to the Underworld, one of the sights is the Stygian Elm:

In medio ramos annosaque bracchia pandit

ulmus opaca, ingens, quam sedem somnia vulgo

uana tenere ferunt, foliisque sub omnibus haerent.

[:Spreads in the midst her boughs and agéd arms

an elm, huge, shadowy, where vain dreams, 'tis said,

are wont to roost them, under every leaf close-clinging.]

Virgil refers to a Roman superstition (vulgo) that elms were trees of ill-omen because their fruit seemed to be of no value.^[61] It has been noted^[62] that two elm-motifs have arisen from classical literature: (1) the 'Paradisal Elm' motif, arising from pastoral idylls and the elm-and-vine theme, and (2) the 'Elm and Death' motif, perhaps arising from Homer's commemorative elms and Virgil's Stygian Elm. Many references to elm in European literature from the Renaissance onwards fit into one or other of these categories.

There are two examples of pteleogenesis (:birth from elms) in world myths. In Germanic and Scandinavian mythology the first woman, Embla, was fashioned from an elm,^[63] while in Japanese mythology Kamuy Fuchi, the chief goddess of the Ainu people, "was born from an elm impregnated by the Possessor of the Heavens".^[64]

The elm occurs frequently in English literature, one of the best known instances being in Shakespeare's A Midsummer Night's Dream, where Titania, Queen of the Fairies, addresses her beloved Nick Bottom using an elm-simile. Here, as often in the elm-and-vine motif, the elm is a masculine symbol:

Sleep thou, and I will wind thee in my arms.

... the female Ivy so

Enrings the barky fingers of the Elm.

O, how I love thee! how I dote on thee!^[65]

Another of the most famous kisses in English literature, that of Paul and Helen at the start of Forster's Howards End, is stolen beneath a great wych elm.

The Elm Tree is also referenced in children's literature. An Elm Tree and Three Sisters is a children's book about three young sisters that plant a small elm tree in their backyard. Throughout the years, the sisters begin to realize that the elm tree becomes an integral part of their lives. The sisters see that the elm tree has been with them through many important milestones through life and they grow old with the tree.

^[66]

The elm in politics[edit]

The Cutting of the elm was a diplomatic altercation between the Kings of France and England in 1188, during which an elm tree near Gisors in Normandy was felled.

In politics the elm is associated with revolutions. In England after the Glorious Revolution of 1688, the final victory of parliamentarians over monarchists, and the arrival from Holland, with William & Mary, of the 'Dutch Elm' hybrid, planting of this cultivar became a fashion among enthusiasts of the new political order.^[67][68]

In the American Revolution 'The Liberty Tree' was an American white elm in Boston, Massachusetts, in front of which, from 1765, the first resistance meetings were held against British attempts to tax the American colonists without democratic representation. When the British, knowing that the tree was a symbol of rebellion, felled it in 1775, the Americans took to widespread 'Liberty Elm' planting, and sewed elm symbols on to their revolutionary flags.^[69][70]Elm-planting by American Presidents later became something of a tradition.

In the French Revolution, too, Les arbres de la liberté (:Liberty Trees), often elms, were planted as symbols of revolutionary hopes, the first in Vienne, Isère, in 1790, by a priest inspired by the Boston elm.^[69] L'Orme de La Madeleine (:the Elm of La Madeleine), Faycelles, Département de Lot, planted around 1790 and surviving to this day, was a case in point.^[71] By contrast, a famous Parisian elm associated with the Ancien Régime, L’Orme de Saint-Gervais in the Place St-Gervais, was felled by the revolutionaries; church authorities planted a new elm in its place in 1846, and an early 20th-century elm stands on the site today.^[72] Premier Lionel Jospin, obliged by tradition to plant a tree in the garden of the Hôtel Matignon, the official residence and workplace of Prime Ministers of France, insisted on planting an elm, so-called 'tree of the Left', choosing the new disease-resistant hybrid 'Clone 762' (Ulmus 'Wanoux' = Vada).^[73] In the French Republican Calendar, in use from 1792 to 1806, the 12th day of the month Ventôse (= 2 March) was officially named «jour de l'Orme», Day of the Elm.

Liberty Elms were also planted in other countries in Europe to celebrate their revolutions, an example being L'Olmo di Montepaone, L'Albero della Libertà(:the Elm of Montepaone, Liberty Tree) in Montepaone, Calabria, planted in 1799 to commemorate the founding of the democratic Parthenopean Republic, and surviving till it was brought down by a recent storm (it has since been cloned and 'replanted').^[74] And after the Greek Revolution of 1821-32, a thousand young elms were brought to Athens from Missolonghi, "Sacred City of the Struggle" against the Turks and scene of Lord Byron's death, and planted in 1839-40 in the National Garden.^[75][76] In an ironic development, wild elms have spread and taken over the grounds of the abandoned Greek royal summer palace at Tatoi in Attica.

Planting a Liberty Tree (un arbre de la liberté) during the French Revolution. Jean-Baptiste Lesueur, 1790

Balcony with elm symbol, overlooking the'Crossroads of the Elm', Place Saint-Gervais, Paris^[72]

President George W. Bush and Laura Bush planting a disease-resistant 'Jefferson' Elmbefore the White House, 2006

Elm suckers spreading before the abandoned summer royal palace in Tatoi, Greece, Μarch 2008

The propagation of elms[edit]

A rooted cutting of European White Elm (July)

Elm propagation methods vary according to elm type and location, and the plantsman’s needs. Native species may be propagated by seed. In their natural setting native species, such as wych elm and European White Elm in central and northern Europe and Field Elmin southern Europe, set viable seed in ‘favourable’ seasons. Optimal conditions occur after a late warm spring.^[1] After pollination, seeds of spring-flowering elms ripen and fall at the start of summer (June); they remain viable for only a few days. They are planted in sandy potting-soil at a depth of one centimetre, and germinate in three weeks. Slow-germinating American Elm will remain dormant until the second season.^[77] Seeds from autumn-flowering elms ripen in the Fall and germinate in the spring.^[77] Since elms may hybridize within and between species, seed-propagation entails a hybridisation risk. In unfavourable seasons elm seeds are usually sterile. Elms outside their natural range, such as Ulmus procera in England, and elms unable to pollinate because pollen-sources are genetically identical, are sterile and are propagated by vegetative reproduction. Vegetative reproduction is also used to produce genetically identical elms (clones). Methods include the winter transplanting of root-suckers; taking hardwood cuttings from vigorous one-year-old shoots in late winter,^[78] taking root-cuttings in early spring; taking softwood cuttings in early summer;^[79] grafting; layering; andmicropropagation. A bottom heat of 18 degrees^[80] and humid conditions are maintained for hard- and softwood cuttings. The transplanting of root-suckers remains the easiest and commonest propagation-method for European Field Elm and its hybrids. For 'specimen' urban elms, grafting to wych-elm root-stock may be used to eliminate suckering or to ensure stronger root-growth. The mutant-elm cultivars are usually grafted, the ‘weeping’ elms 'Camperdown' and 'Horizontalis' at 2 – 3 m, the dwarf cultivars 'Nana' and 'Jacqueline Hillier' at ground level. Since the Siberian Elm is drought-tolerant, in dry countries new varieties of elm are often root-grafted on this species.^[81]

Ripe seeds of Field Elm

Rock elm Ulmus thomasii germinating

Seedling of wych elmUlmus glabra(photograph: Mihailo Grbić)

Root cuttings of Ulmus 'Dodoens' (photograph: Mihailo Grbić)

Rooted hardwood elm-cutting (photograph: Mihailo Grbić)

Rooting of softwood elm-cuttings under mist-propagation system (photograph: Mihailo Grbić)

Mutant variegated Smooth-leafed Elm graft (photograph: Mihailo Grbić)

Root-suckers spreading from Field Elm Ulmus minor

Organisms associated with elm[edit]

'Pouch' leaf-galls on a wych elm (aphidTetraneura ulmi), Germany.

'Pouch' leaf-gall on elm leaf (aphid Tetraneura ulmi), Netherlands.

'Cockscomb' leaf-galls (aphid Colopha compressa), Poland.

'Bladder' leaf-galls on elm leaves (aphidEriosoma lanuginosum), Italy.

'Bladder' leaf-galls on a narrow-leaved elm (aphidEriosoma lanuginosum), Italy.

Aphids in leaf-gall, Poland.

'Pimple' leaf-galls on a field elm (mite Eriophyes ulmi), Spain.

White-letter HairstreakSatyrium w-album, onLutece, Sweden. The larvae feed only on elm.

Egg of Satyrium w-albumnear flower-bud of an elm.

Elm-bark beetle Scolytus multistriatus (size: 2–3 mm), a vector for Dutch Elm Disease

Scolytus multistriatus'galleries' under elm-bark.

Elm-leaf beetleXanthogaleruca luteola, which causes serious damage to elm foliage.

Xanthogaleruca luteola: caterpillar on elm-leaf, Germany.

Elm-leaf damage caused by Xanthogaleruca luteola, Germany.

Bacterial infectionErwinia carotovora of elm sap, which causes 'slime flux' ('wetwood') and staining of the trunk (here on a Camperdown Elm).

Elm

Ulmus minor, East Coker, Somerset, UK.
Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Rosids
Order:	Rosales
Family:	Ulmaceae
Genus:	*Ulmus* L.
Species
See Alphabetical list of extant elm species by scientific name Elm species, varieties, by common name Elm species, varieties, by scientific name Elm cultivars, hybrids and hybrid cultivars Elm synonyms and accepted names

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